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Ramírez, A., & Gutiérrez-Fonseca, P.E. 2020. Freshwater research in Latin America: Current research topics, challenges, and opportunities. Revista de Biología Tropical 68 (Suppl. 2): 1-12. (pdf)

Walteros, J.M., & Ramírez, A. 2020. Urban streams in Latin America: current conditions and research needs. Revista de Biología Tropical 68 (Suppl. 2): 13-28. (pdf)

Ramírez A, Maldonado-Benítez N, Mariani-Ríos A, Figueroa-Santiago J. 2020. Dragonflies and damselflies (Odonata) from Puerto Rico: a checklist with notes on distribution and habitat. PeerJ 8:e9711 (pdf)

Gutiérrez-Fonseca PE, & A. Ramírez. 2020. Mayfly emergence production and body length response to hydrology in a tropical lowland stream. PeerJ 8:e9883 DOI 10.7717/peerj.9883 (pdf)

Boyero L, Pearson RG, Albariño RJ, Callisto M, Correa-Araneda F, Encalada AC, Masese F, Moretti MS, Ramírez A, Sparkman AE, Swan CM. 2020. Identifying stream invertebrates as plant litter consumers. In Baerlocher, F., Gessner, M.O., Graca, M.A.S. (eds). Methods to study litter decomposition - A practical guide. Springer. (link)

Novelo Gutiérrez R, & Ramírez A. 2020. First record of Perithemis thais Kirby, 1889 for Central America (Odonata: Anisoptera: Libellulidae). Argia 32: 29.

Van Beusekom, A.E., G. González, S. Stankavich, J.K. Zimmerman, & A. Ramírez. 2020. Understanding Tropical Forest Abiotic Response to Hurricanes using Experimental Manipulations, Field Observations, and Satellite Data. Biogeosciences 17: 3149–3163. (pdf)

Gutiérrez-Fonseca, P. E., Ramírez, A., Pringle, C. M., Torres, P. J., McDowell, W. H., Covich, A., Todd Crowl, & Pérez-Reyes, O. (2020). When the rainforest dries: Drought effects on a montane tropical stream ecosystem in Puerto Rico. Freshwater Science, 39(2), 197-212. (link)

Rosas, K.G., Colón-Gaud, C. & A. Ramírez, 2020. Trophic basis of production in tropical headwater streams, Puerto Rico: an assessment of the importance of allochthonous resources in fueling food webs. Hydrobiologia 847: 1961–1975. (link)

Peniston, J.H., P.A. Gómez-Ruiz, P. Panwar, H. Uno, & A. Ramírez. 2020. Target species affects the duration of competitive interactions in the Neotropical dragonfly, Micrathyria atra (Odonata: Libellulidae). Notulae odonatologicae 9(5): 173-177.


Classen‐Rodríguez, L, Gutiérrez‐Fonseca, PE, Ramírez, A. 2019. Leaf litter decomposition and macroinvertebrate assemblages along an urban stream gradient in Puerto Rico. Biotropica 51(5): 641-651. DOI.

Kelly, S.P., E. Cuevas, & A. Ramirez. 2019. Urbanization increases the proportion of aquatic insects in the diets of riparian spiders. Freshwater Science 38 (2): 379-390. (pdf)

McDowell, W.H., W.G. McDowell, J.D. Potter, & A. Ramírez. 2019. Nutrient export and elemental stoichiometry in an urban tropical river. Ecological Applications e01839. (enlace / pdf: contact us)

Ríos-Touma, B. & A. Ramírez. 2019. Multiple Stressors in the Neotropical Region: Environmental Impacts in biodiversity hotspots. In: S. Sabater, A. Elosegi, & R. Ludwig (eds). Multiple Stressors in River Ecosystems. Elsevier, Cambridge, MA. (enlace / pdf: contact us)


Novelo-Gutiérrez, R., A. Ramírez, & E. González-Soriano. 2018. Chapter 14.2 - Superfamily Gomphoidea. In: N. Hamada, J. H. Thorp, D.C. Rogers (eds.), Thorp and Covich's Freshwater Invertebrates (Fourth Edition), Pages 377-397, Academic Press. (pdf: contact us)

Román_Heracleo, J., M. Springer, & A. Ramírez. 2018. The larva of Perissolestes remotus (Williamson & Williamson, 1924) (Zygoptera: Perilestidae). International Journal of Odonatology.

Ramírez, A., P.E. Gutiérrez-Fonseca, S.P. Kelly, A.C. Engman, K. Wagner, K.G. Rosas & N. Rodríguez. 2018. Drought Facilitates Species Invasions in an Urban Stream: Results From a Long-Term Study of Tropical Island Fish Assemblage Structure. Frontiers in Ecology and Evolution 6(115).

Gutiérrez-Fonseca, P.E., A. Ramírez, C.M. Pringle. 2018. Large-scale climatic phenomena drive fluctuations in macroinvertebrate assemblages in lowland tropical streams, Costa Rica: The importance of ENSO events in determining long-term (15y) patterns. PLOS ONE 13(2): e0191781. (pdf)

Reyes-Torres, L.J. & A. Ramírez. 2018. Life history and phenology of Phylloicus pulchrus (Trichoptera: Calamoceratidae) in a tropical rainforest stream of Puerto Rico. Revista de Biologia Tropical 66(2): 814-825. (pdf)

Reyes-Torres, L.J. & A. Ramírez. 2018. Effects of experimental pool level reduction on Phylloicus pulchrus (Trichoptera: Calamoceratidae) feeding and cospecific behavior from a tropical Rainforest stream. Intropica 13. 10.21676/23897864.2352 (pdf)


Sanchez, J., A. Ramírez, & S. Kelly. 2017. Decreases in the size of riparian orb webs along an urbanization gradient. Journal of Arachnology 45:248-252.

Boyero, L., M.A.S. Graça, A.M. Tonin, J. Pérez, A.J. Swafford, V. Ferreira, A. Landeira-Dabarca, M.A. Alexandrou, M.O. Gessner, B.G. McKie, R.J. Albariño, L.A. Barmuta, M. Callisto, J. Chará, E. Chauvet, C. Colón-Gaud, D. Dudgeon, A.C. Encalada, R. Figueroa, A.S. Flecker, T. Fleituch, A. Frainer, J.F. Gonçalves Jr., J.E. Helson, T. Iwata, J. Mathooko, C. M’Erimba, C.M. Pringle, A. Ramírez, C.M. Swan, C.M. Yule, & R.G. Pearson. 2017. Riparian plant litter quality increases with latitude. Scientific Reports, 7: 10562. (pdf)

Forero Céspedes, A.M., P.E. Gutiérrez Fonseca, A. Ramírez. 2017. Facilitation between shrimps and mayflies in montane stream ecosystems, Puerto Rico. Rev. Asoc. Col. Cienc. (Col.) 29: 15-20.(pdf: contact us)

Reyes-Maldonado, R., J.A. Sanchez, A. Ramírez and S.P. Kelly. 2017. Riparian spider communities as indicators of stream ecosystem condition in the Río Piedras watershed of Puerto Rico. Actualidades Biológicas 39(107). DOI: 10.17533/udea.acbi.v39n107a07 (pdf)


García, P., R. Novelo-Gutiérrez, G. Vázquez, & A. Ramírez. 2016. Allochthonous vs. autochthonous energy resources for aquatic insects in cloud forest streams, Veracruz, Mexico. Hidrobiológica 26 (3): 483-496. (pdf: contact us)

Reyes-Torres, L.J., Meléndez-Torres, Y. & Ramírez, A. 2016. Occurrence of nematodes on Ephemeroptera nymphs in a tropical rainforest stream. Revista Intropica 11: 67-72. (pdf)

Gutiérrez-Fonseca P. E. & A. Ramírez. 2016. Evaluación de la calidad ecológica de los ríos en Puerto Rico: principales amenazas y herramientas de evaluación. Hidrobiológica 26 (3): 433-441. (pdf: contact us)

Cardona-Rivera, G.A. & A. Ramírez. 2016. Predation of Telebasis vulnerata (Odonata: Coenagrionidae) eggs by detritivorous caddisfly larva, Phylloicus pulchrus (Trichoptera: Calamoceratidae). International Journal of Odonatology 19(4): 253-256. DOI: 10.1080/13887890.2016.1258372. (pdf: contact us)

Wehrtmann, I. S., Ramírez, A., & Pérez-Reyes, O. 2016. Freshwater decapod diversity and conservation in Central America and the Caribbean. In: T. Kawai & N. Cumberlidge (eds.). A Global Overview of the Conservation of Freshwater Decapod Crustaceans. (pp. 267-301): Springer. (pdf: contact us)

Novelo-Gutiérrez, R., A. Ramírez, & D. Delgado. 2016. The larvae of Epigomphus jannyae Belle, 1993 and E. tumefactus Calvert, 1903 (Insecta: Odonata: Gomphidae). PeerJ 4:e2338 (pdf)

Pringle C.M., E.P. Anderson, M. Ardón, R.J. Bixby, S. Connelly, J.H. Duff, A.P. Jackman, P. Paaby, A. Ramírez, G.E. Small, M.N. Snyder, & F.J. Triska. 2016. Rivers of Costa Rica. In: Costa Rican Ecosystems. Ed: M. Kappelle. University of Chicago Press. Chicago, IL. (pdf: contact us)

Ramírez, A. & P.E. Gutiérrez-Fonseca. 2016. Sobre ensambles y ensamblajes ecológicos-respuesta a Monge-Nájera. Revista de Biología Tropical 64(2): 817-819. (pdf)

Boyero, L., R. G. Pearson, C. Hui, M. O. Gessner, J. Pérez, M.A. Alexandrou, M.A.S. Graca, B.J. Cardinale, R.J. Albariño, M. Arunachalam, L.A. Barmuta, A.J. Boulton, A. Bruder, M. Callisto, E. Chauvet, R.G. Death, D. Dudgeon, A.C. Encalada, V. Ferreira, R. Figueroa, A.S. Flecker, J.F. Goncalves Jr, J. Helson, T. Iwata, T. Jinggut, J. Mathooko, C. Mathuriau, C. M’Erimba, M.S. Moretti, C.M. Pringle, A. Ramirez, L. Ratnarajah, J. Rincon, & C.M. Yule. 2016. Biotic and abiotic variables influencing plant litter breakdown in streams: a global study. In Proc. R. Soc. B 283(1829): 20152664. (pdf: contact us)

Capps, K.A. C.N. Bentsen, & A. Ramírez. 2016. Poverty, urbanization, and environmental degradation: urban streams in the developing world. 35(1):429–435. (pdf)

Roy, A.H., K.A. Capps, R.W. El-Sabaawi, K.L. Jones, T.B. Parr, A. Ramírez, R.F. Smith, C.J. Walsh, & S.J. Wenger. 2016. Urbanization and stream ecology: diverse mechanisms of change. Freshwater Science 35(1):272–277. (pdf)

Small, G.E., M. Ardón, J.H. Duff, A.P. Jackman, A. Ramírez, F.J. Triska, & C.M. Pringle. 2016. Phosphorus retention in a lowland Neotropical stream following an eight-year enrichment experiment. Freshwater Science 35(1): 1-11. (pdf)

Astudillo, M.R., R. Novelo-Gutiérrez, G. Vázquez, J.G. García-Franco, & A. Ramírez. 2016. Relationships between land cover, riparian vegetation, stream characteristics, and aquatic insects in cloud forest streams, Mexico. Hydrobiologia 768: 167-181. (pdf: contact us)


Ramírez-Babativa, D., Vázquez, G., Ramírez, A., & Caballero, M. 2015. Dinámica metabólica del periliton de ríos de la cuenca alta del río La Antigua, Veracruz, México. In: J. Alcocer, M. Merino-Ibarra & E. Escobar-Briones (Eds.), Tendencias de investigación en Limnología tropical: Perspectivas universitarias en Latinoamérica.México: Asociación Mexicana de Limnología, A.C., Instituto de Ciencias del Mar y Limnología, UNAM, y Consejo Nacional de Ciencia y Tecnología. (pdf)

Kelly, S.P., E. Cuevas, & A. Ramírez. 2015. Stable isotope analyses of web-spinning spider assemblages along a headwater stream in Puerto Rico. PeerJ 3:e1324; DOI 10.7717/peerj.1324 (pdf)

Gutiérrez-Fonseca, P.E., A.M. Alonso-Rodríguez, A. Cornejo, A.C. Bailey, J. M. Maes & A. Ramírez. 2015. New records of Anacroneuria Klapálek, 1909 (Plecoptera: Perlidae) for Central America. Zootaxa 3994(3):445-448. (pdf: contact us)

Boyero, L., R. G. Pearson, M. O. Gessner, D. Dudgeon, A. Ramírez, C. M. Yule, M. Callisto, C. M. Pringle, A. C. Encalada, M. Arunachalam, J. Mathooko, J. E. Helson, J. Rincón, A. Bruder, A. Cornejo, A. S. Flecker, C. Mathuriau, C. M’Erimba, J. F. G. Jr, M. Moretti11 and T. Jinggu. 2015. Leaf-litter breakdown in tropical streams: is variability the norm? Freshwater Science 34(2): 759-769. (pdf: contact us)

Ramírez, A., M. Ardón, M.M. Douglas, & M.A.S. Graça. 2015. Tropical freshwater sciences: an overview of ongoing tropical research. Freshwater Sciences 34(2): 606-608. (pdf: contact us)


Ramírez, A. & P.E. Gutiérrez-Fonseca. 2014. Puerto Rico. En: Alonso-EguíaLis, P., Mora, J.M., Campbell, B. y M. Springer (eds). Diversidad, conservación y uso de los macroinvertebrados dulceacuícolas de México, Centroamérica, Colombia, Cuba y Puerto Rico. Instituto Mexicano de Tecnología del Agua, Jiutepec, Morelos, México. (pdf)

Muñoz-Erickson, T.A.; Lugo, A.E.; Meléndez-Ackerman, E.; Santiago-Acevedo, L.E.; Seguinot-Barbosa, J.; Méndez-Lázaro, P.; Hall, M.; Quintero, B.; Ramírez, A.; García-Montiel, D.; Pontius Jr., R.G.; Ramos-González, O.M.; Santiago-Bartolomei, R.; Verdejo-Ortíz, J.; Ortíz-Zayas, J.R.; Concepción, C.M.; Cusack, D.; Giusti, J.; McDowell, W.; Cruz-Torres, M.L.; Vallejo, J.; Cray, L.; Zimmerman, J.; Cuadrado-Landrau, V.; & Figueroa, M. 2014. Knowledge to Serve the City: Insights from an Emerging Knowledge-Action Network to Address Vulnerability and Sustainability in San Juan, Puerto Rico. Cities and the Environment 7(1): Article 5. (pdf)

Rodríguez, N. & A. Ramírez. 2014. Protocolo de evaluación visual de quebradas para Puerto Rico. Universidad de Puerto Rico, recinto de Río Piedras. Versión Mayo 2014. (pdf) Mas detalles acá.

Ramírez, A., K.G. Rosas, A. Lugo, O. Ramos-González. 2014. Spatio-temporal variation in stream water chemistry in a tropical urban watershed. Ecology and Society 19(2): 45. (pdf)

Ramírez, A. & P.E. Gutiérrez-Fonseca. 2014. Functional feeding groups of aquatic insect families in Latin America: a critical analysis and review of existing literature. Revista de Biología Tropical 62(Sup.2): 155-167. (pdf)

Torres, P.J. & A. Ramírez. 2014. Land use effects on leaf litter breakdown in low-order streams draining a rapidly developing tropical watershed in Puerto Rico. Revista de Biología Tropical 62(Sup.2): 129-142. (pdf)

Astudillo, M.R., A. Ramírez, R. Novelo-Gutiérrez, & G. Vázquez. 2014. Descomposición de hojarasca en seis arroyos de Bosque Mesófilo de Montaña en la cuenca alta del río La Antigua, Veracruz, México. Revista de Biología Tropical 62(Sup.2): 111-127. (pdf)

Macías, N.A., C. Colón-Gaud, J.W. Duggins, & A. Ramírez. 2014. Do omnivorous shrimp influence mayfly nymph life history traits in a tropical island stream? Revista de Biología Tropical 62(Sup.2): 41-51. (pdf)

Ramírez, A. & P.E. Gutiérrez-Fonseca. 2014. Estudios sobre macroinvertebrados acuáticos en América Latina: avances recientes y direcciones futuras. Revista de Biología Tropical 62(Sup.2): 9-20. (pdf)

Burgos-Caraballo, S., Cantrell, S., & Ramírez, A. 2014. Diversity of benthic biofilms along a land use gradient in tropical headwater streams, Puerto Rico. Microbial Ecology. doi: 10.1007/s00248-014-0401-x. (pdf: contact us)

Gómez-Anaya, J.A., R. Novelo-Gutiérrez, A. Ramírez, and R. Arce-Pérez. 2014. Using empirical field data of aquatic insects to infer a cut-off slope value in asymptotic models to assess inventory completeness. Revista Mexicana de Biodiversidad. 85:218-227. (pdf)


Gutiérrez-Fonseca, P.E., K.G. Rosas, & A. Ramírez. 2013. Aquatic insects of Puerto Rico: a list of families. Dugesiana 20(2): 215-219. (pdf)

Novelo-Gutiérrez, R & A. Ramírez. 2013. Telebasis filiola Perty (Odonata: Coenagrionidae) preying on small web-building spiders (Arachnida: Tetragnathidae). International Journal of Odonatology. DOI: 10.1080/13887890.2013.830992. (pdf: contact us)

Gómez-Anaya, J.A., R. Novelo-Gutiérrez & A. Ramírez. 2013. Temporal variation on Odonata larval assemblage diversity in a lowland stream in western Mexico. Odonatologica 42(4): 309-323. (pdf)

Ramírez A. & P.E. Gutiérrez-Fonseca. 2013. The larvae of Heteragrion majus Selys and H. atrolineatum Donnelly, with a key to known species from Costa Rica (Odonata: Megapodagrionidae). Zootaxa 3609 (1): 096–100. (pdf: contact us)

Ardón, M., J.H. Duff, A. Ramírez, G.E. Small, A.P. Jackman, F.J. Triska & C.M. Pringle. 2013. Experimental acidification of two biogeochemically-distinct neotropical streams: buffering mechanisms and macroinvertebrate drift. Science of the Total Environment 443: 267–277. (pdf)

Gutiérrez-Fonseca, P.E., A. Ramírez, G. Umaña & M. Springer. 2013. Macroinvertebrados dulceacuícolas de la Isla del Coco, Costa Rica: especies y comparación con otras islas del Pacífico Tropical Oriental. Revista de Biología Tropical. 61(2):657-668. (pdf)


Small, G.E., Ardón, M., Jackman, A.P., Duff, J.H., Triska, F.J., Ramirez, A., Snyder, M., Pringle, C.M. 2012. Rainfall-driven amplification of seasonal acidification in poorly buffered tropical streams. Ecosystems 15:974-985. (pdf)

Engman, A.C. and A. Ramírez. 2012. Fish assemblage structure in urban streams of Puerto Rico: The importance of reach- and catchment-scale abiotic factors. Hydrobiologia 693: 141-155. (pdf)

Ramírez, A., A. Engman, K.G. Rosas, O. Perez-Reyes, and D.M. Martinó-Cardona. 2012. Urban impacts on tropical island streams: some key aspects influencing ecosystem response. Urban Ecosystems 15(2): 315-325. (pdf)

Boyero L., R.G. Pearson, D. Dudgeon, V. Ferreira, M.A.S. Graça, M.O. Gessner, A.J. Boulton, E. Chauvet, C.M. Yule, R.J. Albariño, A. Ramírez, J.E. Helson, M. Callisto, M. Arunachalam, J. Chará, R. Figueroa, J.M. Mathooko, J.F. Gonçalves Jr., M. Moretti, A.M. Chará, J.N. Davies, A. Encalada, S. Lamothe, L.M. Buria, J. Castela, A. Cornejo, A.O.Y. Li, C. M’Erimba, V.D. Villanueva, M.C. Zúñiga, C.Swan & L.A. Barmuta. 2012. Global patterns of distribution in stream detritivores: implications for biodiversity loss in changing climates. Global Ecology and Biogeography 21:134-141. (pdf)


Ramírez, A., M. Altamiranda-Saavedra, P. Gutiérrez-Fonseca & M. Springer. 2011. The neotropical damselfly genus Cora: New larval descriptions and a comparative analysis of larvae of known species (Odonata: Polythoridae). International Journal of Odonatology 14(3): 249–256. (pdf)

Boyero, L., R. Pearson, D. Dudgeon, M. Graca, M. Gessner, R. Albarino, V. Ferreira, C. Yule, A. Boulton, M. Arunachalam, M. Callisto, E. Chauvet, A. Ramirez, J. Chara, M. Moretti, J. Goncalves Jr, J. Helson, A.M. Chara, A. Encalada, J. Davies, S. Lamothe, A. Cornejo, J. Castela, A. Li, L. Buria, V. Díaz Villanueva, M.C. Zuniga, & C. Pringle. 2011. Global distribution of a key trophic guild contrasts with common latitudinal diversity patterns. Ecology 92: 1839-1848. (pdf)

Boyero, L., R.G. Pearson, M.O. Gessner, L.A. Barmuta, V. Ferreira, M.A.S. Graça, D. Dudgeon, A.J. Boulton, M. Callisto, E. Chauvet, J.E. Helson, A. Bruder, R.J. Albariño, C.M. Yule, M. Arunachalam, J.N. Davies, R. Figueroa, A.S. Flecker, A. Ramírez, R.G. Death, T. Iwata, J.M. Mathooko, C. Mathuriau, J.F. Gonçalves Jr., M.S. Moretti, T. Jingut, S. Lamothe, C. M’Erimba, L. Ratnarajah, M.H. Schindler, J. Castela, L.M. Buria, A. Cornejo, V.D. Villanueva, & D.C. West. 2011. A global experiment suggests climate warming will not accelerate litter decomposition in streams but may reduce carbon sequestration. Ecology Letters 14:289-294. (pdf)

De Jesús-Crespo, R. & A. Ramírez. 2011. Effects of urbanization on stream physicochemistry and macroinvertebrate assemblages in a tropical urban watershed in Puerto Rico. Journal of the North American Benthological Society 30(3):739-750. (pdf)

De Jesús-Crespo, R. & A. Ramírez. 2011. The use of a stream visual assessment protocol to determine ecosystem integrity in an urban watershed in Puerto Rico. Physics and Chemistry of the Earth 36(12):560-566. (pdf)


Springer, M., A. Ramírez, & P. Hanson. 2010. Macroinvertebrados de Agua Dulce de Costa Rica I. Revista de Biología Tropical 58 – Supplement No. 4. (web)

Hanson, P., M. Springer, & A. Ramírez. 2010. Introducción a los grupos de macroinvertebrados acuáticos. Revista de Biología Tropical 58(Suppl. 4): 3-37. (pdf)

Ramírez, A. 2010. Métodos de recolección. Revista de Biología Tropical 58(Suppl. 4): 41-50. (pdf)

Ramírez, A. 2010. Odonata. Revista de Biologia Tropical 58(Suppl. 4): 97-136. (pdf. Versiones actualizadas en nuestra página)

Benstead, J.P, W.F. Cross, J.G. March, W.H. McDowell, A. Ramírez, & A. Covich. 2010. Biotic and abiotic controls on ecosystem significance of consumer excretion in two contrasting tropical streams. Freshwater Biology 55: 2047-2061.


Blanco, J.F., A. Ramírez, & F.N. Scatena. 2009. The streams of Gorgona Natural National Park within the global context: An introduction to the special issue. Actualidades Biologicas 31(91): 105-110. (pdf)

Ramírez, A., R. De Jesús-Crespo, D.M. Martinó-Cardona, N. Martinez-Rivera, & S. Burgos-Caraballo. 2009. Urban streams in Puerto Rico: what can we learn from the tropics? Journal of the North American Benthological Society 28(4): 1070-1079. (pdf)

Wenger, S.J., A.H. Roy, C.R. Jackson, E.S. Bernhardt, T.L. Carter, S. Filoso, C.A. Gibson, W.C. Hession, S.S. Kausha, E. Martı, J.L. Meyer, M.A. Palmer, M.J. Paul, A.H. Purcell, A. Ramirez, A.D. Rosemond, K.A. Schofield, E.B. Sudduth & C.J. Walsh. 2009. Twenty-six key research questions in urban stream ecology: an assessment of the state of the science. Journal of the North American Benthological Society 28(4): 1080-1098. (pdf)

Boyero, L., A. Ramírez, D. Dudgeon, & R.G. Pearson. 2009. Are tropical streams really different? Journal of the North American Benthological Society 28(2): 397-403. (pdf)


Cross, W.F., A.P. Covich, T.A. Crowl, J.P. Benstead, & A. Ramírez. 2008. Secondary production, longevity and resource consumption rates of freshwater shrimps in two tropical streams with contrasting geomorphology and food web structure. Freshwater Biology DOI:10.1111/j.1365-2427.2008.02078.x (pdf)

Small, G., C.M. Pringle, F.J. Triska, J.H. Duff, A.P. Jackman, M. Hidalgo, A. Ramírez, & M. Ardón. 2008. The dynamics of phosphorus retention during an eight-year P-addition in a neotropical headwater stream. Verh. Internat.Verein. Limnol. 30: 551-554. (pdf)

Cross, W.F., A. Ramírez, A. Santana, & L. Silvestrini. 2008. Toward separating the relative importance of invertebrate consumption and bioturbation in Puerto Rican streams. Biotropica. 40(4): 477–484. (pdf)

Ramírez, A., C.M. Pringle, & K.M. Wantzen. 2008. Tropical river conservation. In D. Dudgeon (ed.), Tropical Stream Ecology. Academic Press. (pdf)


Ramírez, A. 2007. Biodiversidad de insectos acuáticos y el funcionamiento de los ecosistemas. En: Entomología Acuática Mexicana: estado actual de conocimiento y aplicación. ISBN: 978-968-5536-89-9. (pdf: contact us)

Bobeldyk, A. & A. Ramírez. 2007. Leaf breakdown in a tropical headwater stream (Puerto Rico): The role of freshwater shrimps and detritivorous insects. Journal of Freshwater Ecology 22:581-590. (pdf)


Triska, F.J., C.M. Pringle, J.H. Duff, R.J. Avanzino, A. Ramírez, M. Ardon & A.P. Jackman. 2006. Soluble reactive phosphorus transport and retention in tropical, rainforest streams draining a volcanic and geothermally active landscape in Costa Rica. 1: Long-term concentration patterns, pore water environment and response to ENSO events. Biogeochemistry DOI 10.1007/s10533-006-9026-6. (pdf)

Wantzen, K. M., A. Ramírez, & K. Winemiller. 2006. New Vistas in Neotropical Stream Ecology: Introduction. Journal of North American Benthological Society 25:61-65. (pdf)

Ramírez, A., C.M. Pringle, & M. Douglas. 2006. Physico-chemical variables and the structure of stream insect assemblages in a tropical landscape. Journal of North American Benthological Society 25:108-123. (pdf)

Ramírez, A. & C.M. Pringle. 2006. Fast growth and turnover of chironomid assemblages in response to stream phosphorus levels in a tropical lowland landscape. Limnology and Oceanography 51: 189-196. (pdf)


Ramírez, A. & L.R. Hernández-Cruz. 2004. Aquatic insect assemblages in shrimp-dominated tropical streams, Puerto Rico. Biotropica 36: 259-266. (pdf)

Ramírez, A. & C.M. Pringle. 2004. Do macroconsumers affect insect responses to a natural stream phosphorus gradient? Hydrobiologia 515: 235-246. (pdf: contact us)

Ramírez, A. 2004. Ecological Research and the Costa Rican Park System. Ecological Applications 14: 25-27. (pdf)


Ramírez, A., C.M. Pringle, & L. Molina. 2003. Effects of stream phosphorus levels on microbial respiration. Freshwater Biology 48:88-97. (pdf)


Rosemond, A.D., C.M. Pringle, A. Ramírez, M.J. Paul, & J. Meyer. 2002. Landscape variation in phosphorus concentration and effects on detritus-based tropical streams. Limnology and Oceanography 47: 278-289. (pdf)


Rosemond, A.D., C.M. Pringle, A. Ramírez & M.J. Paul. 2001. A test of top-down and bottom-up control in a detritus-based food web. Ecology 82: 2279-2293. (pdf)

Ramírez, A. & C.M. Pringle. 2001. Spatial and temporal patterns of invertebrate drift in streams draining a Neotropical landscape. Freshwater Biology. 46:47-62. (pdf: contact us)


Ramírez, A., D.R. Paulson & C. Esquivel. 2000. Odonata of Costa Rica: Diversity and checklist of species. Revista de Biología Tropical 48(1): 245-252. (pdf)

Ramírez, A. 2000. Dragonflies and damselflies of Costa Rican Cloud Forests, p. 97. In: N. Nadkarni & N.T. Wheelwright (Eds.). Monteverde: Ecology and Conservation of a Tropical Cloud Forest. Oxford University Press, New York. (pdf: contact us)


Ramírez, A. & R. Novelo-Gutiérrez. 1999. The neotropical dragonfly genus Macrothemis: new larval descriptions and an evaluation of its generic status based on larval stages (Odonata: Libellulidae). Journal of the North American Benthological Society 18:66-72. (pdf)


Ramírez, A., P. Paaby, C.M. Pringle & G. Agüero. 1998. Effect of habitat type on benthic macroinvertebrates in two tropical lowland streams, Costa Rica. Revista de Biología Tropical 46 (Suppl. 6): 201-213. (pdf)

Paaby, P., A. Ramírez, & C.M. Pringle. 1998. The benthic macroinvertebrate community in Caribbean Costa Rican streams and the effect of sampling methods. Revista de Biología Tropical 46 (Suppl. 6): 185-199. (pdf)

Ramírez, A. & C.M. Pringle. 1998. Invertebrate drift and benthic community dynamics in a lowland neotropical stream, Costa Rica. Hydrobiologia 386: 19-26. (pdf)

Ramírez, A. & C.M. Pringle. 1998. Structure and production of a benthic insect assemblage in a neotropical stream. Journal of North American Benthological Society 17: 443-463. (pdf)

Novelo-Gutiérrez, R. & A. Ramírez. 1998. The larva of Macrothemis inacuta (Odonata: Libellulidae). Entomological News 109: 301-306. (pdf)

Rosemond, A.D., C.M. Pringle, & A. Ramírez. 1998. Macroconsumer effects on insect detritivores and detritus processing in a tropical stream. Freshwater Biology 39: 515-524. (pdf: contact us)

Pringle, C.M. & A. Ramírez. 1998. Use of both benthic and drift sampling techniques to assess tropical stream invertebrate communities along an altitudinal gradient, Costa Rica. Freshwater Biology 39: 359-373. (pdf)


Ramírez, A. 1997. Lista de especies costarricenses del orden Odonata (Insecta) de las que se conoce la náyade. Revista de Biología Tropical 44/45: 225-232. (pdf)

Ramírez, A. 1997. Description and natural history of the Costa Rican Odonata larvae. V: Megaloprepus caerulatus (Drury, 1782) (Zygoptera, Pseudostigmatidae). Odonatologica 26: 75-81. (pdf)


Ramírez, A. 1996. Six new dragonfly larvae of the family Gomphidae in Costa Rica, with a key to the Central American genera (Anisoptera). Odonatologica 25: 145-158. (pdf)


Novelo-Gutiérrez, R. & A. Ramírez. 1995. The larvae of Neocordulia batesi longipollex Calvert, 1909 (Odonata: Cordulidae). Journal of the New York Entomological Society 103: 180-184. (pdf)

Ramírez, A. 1995. Descripción e historia natural de las larvas de odonatos de Costa Rica. IV: Mecistogaster ornata (Rambur, 1842) (Zygoptera, Pseudostigmatidae). Bulletin of American Odonatology 3: 43-47. (pdf)

De la Rosa C. & A. Ramírez. 1995. A note on phototactic behavior and on phoretic associations in larvae of Mecistogaster ornata Rambur from northern Costa Rica (Pseudostigmatidae, Zygoptera). Odonatologica 24: 219-224. (pdf)


Ramírez-Ulate, A. & R. Novelo-Gutiérrez. 1994. Megapodagrionidae (Odonata, Zygoptera) de México y Centro América I: Las náyades de Philogenia carrillica, P. peacocki y P. terraba. Acta Zoológica Mexicana 63: 61-73. (pdf)

Ramírez, A. 1994. Descripción e historia natural de las larvas de odonatos de Costa Rica. III: Gynacantha tibiata (Karsch, 1891) (Anisoptera; Aeshnidae). Bulletin of American Odonatology 2: 9-14. (pdf)

Ramírez, A. 1994. Descripción e historia natural de las náyades de odonatos de Costa Rica. II: Archilestes neblina (Garrison,1982) (Odonata, Lestidae), con una clave para las especies del género en Costa Rica. Folia Entomológica Mexicana 90: 9-16. (pdf)


Ramírez, A. 1992. Description and natural history of Costa Rican dragonfly larvae. I: Heteragrion erythrogastrum Selys, 1886 (Zygoptera: Megapodagrionidae). Odonatologica 21: 361-365. (pdf)